I think a couple of factors that are often overlooked and misunderstood in boxing are:
1. Boxing stamina is sport specific and "different" than other types of stamina. An example is when Sweet Pea Whitaker fought Gary Jacobs. Jacobs was ripped and in phenomenal shape, known for regularly competing in triathlons/races and running sub 5 minute miles. The guy obviously had great stamina, but in the ring vs the much smaller Whitaker, he couldn't match Pernell's pace or tempo and became exhausted by the end.
2. All the training and natural gifts in the world won't matter if you let your mind/stress/anxiety get the better of you. From my experience, just the pressure of performing in front of a large audience can put pressure on you, cause anxiety, and rob you of your stamina. Staying relaxed is a challenge, and you have to remain calm to have good stamina. Look at JCC as an example, he never got worked up or hyped, he went out like it was just another day and applied steady pressure throughout the fight. It's why/how James Toney could be fat and win a twelve round fight. He was relaxed in the ring and this allowed him to conserve energy and fight at a controlled pace.
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