Re: Which knuckles do the hitting in boxing?
If you feel any trouble in your smaller knuckles when punching, first make sure your hands are wrapped properly in that your hand-wraps do not impede your ability to make a good solid fist (i.e. too much bandage over your palms). The same applies with gloves, make sure you can make a proper fist with them on. Wrapping your fingers around the bar of the glove will help too, as well as to ensure that the small metacarpals in your hand won't shift around when you punch. Then assuming you're throwing your punches properly, and aren't coping a hand injury, the three-knuckle landing as advised by Dempsey should be fine.
Landing with the other knuckles can depend. Say for instance with a long overhand right, the larger knuckles may be better suited, since the force may not be distributed equally if attempted to land with the three knuckles instead. The same maybe true with a jab thrown in an arc to get around a guard.
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